Culture was once considered the patented property of human beings: We have the art, science, music and online shopping; animals have the instinct, imprinting and hard-wired responses. But that dismissive attitude toward nonhuman minds turns out to be more deeply misguided with every new finding of animal wit or whimsy: Culture, as many biologists now understand it, is much bigger than we are. “If you define culture as a set of behaviors shared by a group and transmitted through the group by social learning, then you find that it’s widespread in the animal kingdom,” said Andrew Whiten, a psychologist and neuroscientist at the University of St. Andrews, in Scotland, and the author of the Science review. “You see it from primates and cetaceans, to birds and fish, and now we even find it in insects.”